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B.Sc, Biochemistry and Microbiology, University of Cape Town, Cape Town, South Africa, 1990 B.Sc(MED)(HONS), Medical Biochemistry, University of Cape Town, Cape Town, South Africa, 1991 Ph.D, Medical Biochemistry, University of Cape Town, Cape Town, South Africa, 2000 Postdoctoral Fellowship, National Cancer Institute, National Institutes of Health, Bestheda MD, 2006

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