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研究领域

A number of research projects are currently underway. Several of these are externally funded and all involve student participants. Thermal biology of small lizards from cold climates Temperature-dependent performance of atypical geckos (diurnal and cold-climate species) Evolution and biological consequences of diet, especially herbivory in reptiles Evolution and ecophysiological implications of reproductive modes in squamate reptiles Speciation, species boundaries, and phylogeny of Liolaemus and Phymaturus lizards I enjoy working with undergraduate and graduate students who share my interests in herpetology, or aspects of the evolution, functional morphology, physiology, and ecology of most vertebrates. Our laboratory facilities--coupled with those in the department, university, and the southern California environment--provide excellent opportunities for research. I offer a supportive yet critical environment for fulfilling intellectual and academic pursuits. You can learn more about research opportunities and expectations for investigators in LICH here.

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Espinoza, R. E. 2009. Western fence lizard (Sceloporus occidentalis). Pp. 234-237, in: Lizards of the American Southwest: A Photographic Fieldguide, L. L. C. Jones and R. E. Lovich (eds.). Rio Nuevo Press, Tucson, Arizona. Espinoza, R. E. 2009. Mohave fringe-toed lizard (Uma scoparia). Pp. 278-281, in: Lizards of the American Southwest: A Photographic Fieldguide, L. L. C. Jones and R. E. Lovich (eds.). Rio Nuevo Press, Tucson, Arizona. Espinoza, R. E. 2009. Western banded gecko (Coleonyx variegatus). Pp. 312-315, in: Lizards of the American Southwest: A Photographic Fieldguide. L. L. C. Jones and R. E. Lovich (eds.). Rio Nuevo Press, Tucson, Arizona. Espinoza, R. E. and K. de Queiroz. 2008. Historical perspectives: Richard Emmett Etheridge. Copeia 2008:708-717. [pdf] Abadala, C. S., A. S. Quinteros, and R. E. Espinoza. 2008. Two new species of Liolaemus (Iguania: Liolaemidae) from the Puna of northwestern Argentina. Herpetologica 64:458-471. [pdf] Espinoza, R. E. and S. Quinteros. 2008. A hot knot of toads: aggregation provides thermal benefits to metamorphic Andean toads. Journal of Thermal Biology 33:67-75. [pdf] Tracy, C. R. , K. E. Nussear, T. C. Esque, K. Dean-Bradley, C. R. Tracy, L. A. DeFalco, K. T. Castle, L. C. Zimmerman, R. E. Espinoza, and A. M. Barber. 2006. The importance of physiological ecology in conservation biology. Integrative and Comparative Biology 46:1191-1205. [pdf] Lancaster, J. R., P. Wilson, and R. E. Espinoza. 2006. Physiological benefits as precursors of sociality: why banded geckos band. Animal Behaviour 72:199-207. [pdf] Cruz, F. B., L. A. Fitzgerald, R. E. Espinoza, and Schulte, J. A. II. 2005. The importance of phylogenetic scale in tests of Bergmann's and Rapoport's rules: lessons from a clade of South American lizards. Journal of Evolutionary Biology 18:1559-1574. [pdf] Tracy, C. R. , K. M. Flack, L. C. Zimmerman, R. E. Espinoza, and C. R. Tracy. 2005. Herbivory imposes constraints on voluntary hypothermia in lizards. Copeia 2005:12-19. [pdf] Espinoza, R. E., J. J. Wiens, and C. R. Tracy. 2004. Recurrent evolution of herbivory in small, cold-climate lizards: breaking the ecophysiological rules of reptilian herbivory. Proceedings of the National Academy of Sciences, USA 101:16819-16824. [pdf] Lobo, F. and R. E. Espinoza. 2004. Two new Liolaemus from the Puna region of Argentina and Chile: further resolution of purported reproductive bimodality in Liolaemus alticolor (Iguania: Liolaemidae). Copeia 2004:850-866. [pdf] Hernandez, R. A. and R. E. Espinoza. 2004. Description of the female of the enigmatic lizard, Liolaemus heliodermis (Iguania: Liolaemidae). Herpetological Review 35:227-229. [pdf] Espinoza, R. E. and F. Lobo. 2003. Two new species of Liolaemus lizards from northwestern Argentina: speciation within the northern subclade of the elongatus group (Iguania: Liolaemidae). Herpetologica 59:89-105. [pdf] Cooper, W. E. Jr., J. J. Habegger, and R. E. Espinoza. 2001. Responses to prey and plant chemicals by three iguanian lizards: relationship to plants in the diet. Amphibia-Reptilia 22:349-362. Scheltinga, D. M., B. G. M. Jamieson, R. E. Espinoza, and K. S. Orrell. 2001. Descriptions of the mature spermatozoa of the lizards Crotaphytus bicinctores, Gambelia wislizenii (Crotaphytidae), and Anolis carolinensis (Polychrotidae) (Reptilia, Squamata, Iguania). Journal of Morphology 247:160-171. [pdf] Simandle, E. T., R. E. Espinoza, K. E. Nussear, and C. R. Tracy. 2001. Lizards, lipids, and dietary links to animal function. Physiological and Biochemical Zoology 74:625-640. [pdf] Etheridge, R. and R. E. Espinoza. 2000. Taxonomy of the Liolaeminae (Squamata: Iguania: Tropiduridae) and a semi-annotated bibliography. Smithsonian Herpetological Information Service 126:1-64. [continuously updated on the web: Liolaemidae Website] Espinoza, R. E., F. Lobo, and F. B. Cruz. 2000. Liolaemus heliodermis, a new lizard from northwestern Argentina with remarks on the content of the elongatus group (Iguania: Tropiduridae). Herpetologica 56:235-244.[pdf] Schulte, J. A. II, J. R. Macey, R. E. Espinoza, and A. Larson. 2000. Phylogenetic relationships in the iguanid lizard genus Liolaemus: multiple origins of viviparous reproduction and evidence for recurring Andean vicariance and dispersal. Biological Journal of the Linnean Society 69:75-102. [pdf] Halloy, M. and R. E. Espinoza. 1999-2000. Territorial encounters and threat displays in the Neotropical frog, Phyllomedusa sauvagii (Anura: Hylidae). Herpetological Natural History 7:175-180. [pdf] Lobo, F. and R. E. Espinoza. 1999. Two new cryptic species of Liolaemus (Iguania: Tropiduridae) from northwestern Argentina: resolution of the purported reproductive bimodality of Liolaemus alticolor. Copeia 1999:122-140.[pdf] Nussear, K. E., R. E. Espinoza, C. M. Gubbins, K. J. Field, and J. P. Hayes. 1998. Diet quality does not affect resting metabolic rate or body temperatures selected by an herbivorous lizard. Journal of Comparative Physiology B 168:183-189. [pdf/erratum] Espinoza, R. E. and C. R. Tracy. 1997. Thermal biology, metabolism, and hibernation. Pp. 149-184, in: The Biology, Husbandry and Health Care of Reptiles. L. J. Ackerman (Ed.). Volume 1, Biology of Reptiles. T.F.H. Publications, Neptune City, New Jersey. [pdf] Espinoza, R. E. and F. Lobo. 1996. Possible communal nesting in two species of Liolaemus lizards (Iguania: Tropiduridae) from northern Argentina. Herpetological Natural History 4:65-68. [pdf]

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