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Earl S. Tupper Fellow, Smithsonian Tropical Research Institute, Panama, 2015 Sibbald Fellow, Royal Botanic Garden, Edinburgh, Scotland, 2015 Formation postdoctorale. Institute of Systematic Botany-Munich Botanic Garden, l'Allemagne, 2014. Ph. D. en biologie évolutive, University of Connecticut, USA, 2011. M. Sc. en biologie des plantes, Southern Illinois University, USA, 2005. B. Sc. en biologie, Universidad de Panama, Panama, 2003

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Villarreal, J.C., Crandall-Stotler, B.J., Hart, M.L., Long, D.G. & Forrest, L.L. 2015. Divergence times and the evolution of morphological complexity in an early land plant lineage (Marchantiopsida) with a slow molecular rate. New Phytologist. doi: 10.1111/nph.13716 Villarreal, J.C. & K. Renzaglia. 2015. The hornworts: important advancements in early land plant evolution. Journal of Bryology 37(3): 157-170 (invited review). Brown, R.C., B.E. Lemmon, M. Shimamura, J.C. Villarreal & K.S. Renzaglia. 2015. Spores of relictual bryophytes: diverse adaptations to life on land. Review of Palaeobotany and Palinology 216: 1-17. Wicket, N., --42 authors, D.W. Stevenson, J.C. Villarreal, J.A. Leebens-Mack. 2014. A phylotranscriptomics analysis of the origin and early diversification of land plants. Proceedings of the National Academy of Sciences, USA. Li, F.-W., J.C. Villarreal, --32 authors, D.W. Stevenson, K.M. Pryer. 2014. Horizontal gene transfer of a chimeric photoreceptor, neochrome, from bryophytes to ferns. Proceedings of the National Academy of Sciences, USA 111: 6672-6677,. Villarreal, J.C. & S.S. Renner. 2013. Correlates of monoicy and dioicy in hornworts, the apparent sister group to vascular plants. BMC Evolutionary Biology 13: 239. doi:10.1186/1471-2148-13-239. Villarreal, J.C. & K.S. Renzaglia. 2006. Structure and development of Nostoc strands in Leiosporoceros dussii (Anthocerotophyta): a novel symbiosis in land plants. American Journal of Botany 93: 693-705.

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