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Teaching Fellow in Anatomy, School of Medical Education, Newcastle University (2012-2015) Academic teaching and scholarship position involving lecturing and delivering practical sessions in gross anatomy, histology and embryology for MBBS, BMS and BDS degree programmes, pedagogic research in anatomical and medical education. Senior Postdoctoral Research Associate, Institute of Genetic Medicine, Newcastle University (2010-2012) British Heart Foundation funded postdoctoral research fellowship investigating the role of Wnt planar cell polarity signalling and Rac1 in the regulation of vertebrate cardiovascular development. Postdoctoral Research Fellow, University of St Andrews (2007-2008) Industrially funded research project investigating the regulation of gene expression by insulin-like growth factor signalling in muscle growth. Postdoctoral Research Fellow, University of Dundee (2004-2007) Cancer Research UK fellowship investigating fibroblast growth factor and mitogen-activated protein kinase signalling in the regulation of gene expression in vertebrate development and cancer. PhD Student, University of York (2000-2003) BBSRC studentship involving studying the regulation of Cdx gene expression by fibroblast growth factor and Wnt signalling in head to tail patterning during vertebrate development.

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Backhouse M, Fitzpatrick M, Hutchinson J, Thandi CS, Keenan ID. Improvements in anatomy knowledge when utilizing a novel cyclical “Observe-Reflect-Draw-Edit-Repeat” learning process. Anatomical Sciences Education 2017, 10(1), 7-22. Backhouse M, Fitzpatrick M, Selwyn-Gotha J, Allen R, Keenan I. ORDER (observe-reflect-draw-edit-repeat): a novel cyclical learning process in anatomy education. In: Anatomical Society Summer Meeting: Skin and Bones. 2015, University of Bradford: Wiley-Blackwell Publishing Ltd. Keenan ID, Backhouse M, Bashir A, Allen R. The future ORDER: a student partner approach forevidence-based evaluation and implementation of an ‘observe–reflect–draw–edit–repeat’ artistic learning process in anatomyeducation. In: Anatomical Society Winter Meeting 2013: Assessment Within Medical Education & Sense Organs. 2014, Newcastle upon Tyne, UK: Journal of Anatomy. Backhouse M, Bashir A, Allen R, Jackson S, Keenan ID. The future of anatomy: a student partner approach to theevaluation and implementation of evidence based artisticlearning techniques. In: Anatomical Society Winter Meeting 2013. 2014, Newcastle University, Newcastle upon Tyne, UK: Wiley-Blackwell Publishing Ltd. Matthan J, Keenan I. Acquisition, development and retention of clinical anatomical knowledge by foundation doctors during anatomy demonstrator rotations. In: Anatomical Society Summer Meeting Skin and Bones. 2014, University of Bradford: Wiley-Blackwell Publishing Ltd. Boczonadi V, Gillespie R, Keenan ID, Ramsbottom SA, Donald-Wilson C, Al Nazer M, Humbert P, Schwarz RJ, Chaudhry B, Henderson DJ. Scrib:Rac1 interactions are required for the morphogenesis of the ventricular myocardium. Cardiovascular Research 2014, 104(1), 103-115. Pergolizzi C, Matthan J, Keenan I. Is dissection of more educational value than other methods of learning and teaching anatomy? A student partner approach for evidence-based investigation of dissection as a teaching and learning method in anatomy. In: Anatomical Society Winter Meeting 2013: Assessment Within Medical Education & Sense Organs. 2013, Newcastle University, Newcastle upon Tyne, UK: Wiley-Blackwell Publishing Ltd. Backhouse M, Pergolizzi C, Fitzpatrick M, Allen R, Holmes E, Jackson S, Matthan J, Keenan ID. The future of anatomy education: investigating the impact of student partner approaches for evidence-based evaluation of teaching and learning techniques. In: Anatomical Society Winter Meeting 2013. 2014, Newcastle University, Newcastle upon Tyne, UK: Wiley-Blackwell Publishing Ltd. Keenan ID, Rhee HJ, Chaudhry B, Henderson DJ. Origin of non-cardiac endothelial cells from an Isl1+ lineage. FEBS Letters 2012, 586(13), 1790-1794. Ekerot M, Stavridis MP, Delavaine L, Mitchell MP, Staples C, Owens DM, Keenan ID, Dickinson RJ, Storey KG, Keyse SM. Negative-feedback regulation of FGF signalling by DUSP6/MKP-3 is driven by ERK1/2 and mediated by Ets factor binding to a conserved site within the DUSP6/MKP-3 gene promoter. Biochemical Journal 2008, 412(2), 287-98. Keenan ID, Sharrard RM, Isaacs HV. FGF signal transduction and the regulation of Cdx gene expression. Developmental Biology 2006, 299(2), 478-88. Smith TG, Karlsson M, Lunn JS, Eblaghie MC, Keenan ID, Farrell ER, Tickle C, Storey KG, Keyse SM. Negative feedback predominates over cross-regulation to control ERK MAPK activity in response to FGF signalling in embryos. FEBS Letters 2006, 580(17), 4242-4245. Reece-Hoyes JS, Keenan ID, Pownall ME, Isaacs HV. A consensus Oct1 binding site is required for the activity of the Xenopus Cdx4 promoter. Developmental Biology 2005, 282(2), 509-523. Reece-Hoyes JS, Keenan ID, Isaacs HV. Cloning and expression of the Cdx family from the frog Xenopus tropicalis. Developmental Dynamics 2002, 223(1), 134-140. Hennessy C, Keenan ID, Border S. Anatomy of a tweet: how to use social media as anacademic tool. In: Anatomical Society Summer Meeting at Brighton and Sussex Medical School. 2017, Brighton and Sussex Medical School: Journal of Anatomy. Keenan ID, Backhouse M, Fitzpatrick M, Hutchinson J, Thandi CS. Improvements in anatomy knowledge when using a novel cyclical artistic learning process. In: Anatomical Society Summer Meeting at Brighton and Sussex Medical School. 2017. Woodward R, O'Sullivan J, Holmes E, Keenan ID. Investigating creative writing as an anatomy learning tool. In: Anatomical Society Summer Meeting at Brighton and Sussex Medical School. 2017, Brighton and Sussex Medical School: Journal of Anatomy.

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