This study presents novel findings that expand our understanding of the evolutionary history and diversity of the Limoniidae subfamily Limnophilinae, with a specific focus on the relict genus Chilelimnophila. The family Limoniidae is well documented in the fossil record, with many species considered relicts in the modern fauna. Our study is based on the discovery of the first fossil specimens of Chilelimnophila in both Cretaceous Spanish and Kachin amber. The new materials have allowed the establishment of a new subgenus Chilelimnophila (Ribeironia) Krzemiński, Kania-Kłosok et Arillo, subgen. nov. and three new species: Chilelimnophila (Ribeironia) amorimi Krzemiński, Kania-Kłosok et Arillo, sp. nov., Chilelimnophila (Chilelimnophila) wangi Krzemiński, Kania-Kłosok et Arillo, sp. nov., and Chilelimnophila (Chilelimnophila) parva Krzemiński, Kania-Kłosok et Arillo, sp. nov. This research also significantly extends the stratigraphic range of the genus, providing evidence of its existence dating back to the Early Cretaceous period. The morphology and phylogenetic analysis of Chilelimnophila and related genera within the Limoniidae subfamily Limnophilinae provide valuable insights into the evolutionary relationships of these organisms. Furthermore, our research highlights the unique environmental conditions of Cretaceous Spanish and Kachin amber formations, which differ significantly from the current habitat of these organisms.

中文翻译：

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中生代孑遗属Chilelimnophila（双翅目、柠檬科）的进化史和生物地理分布


这项研究提出了新的发现，扩大了我们对 Limoniidae 亚科 Limnophilinae 的进化历史和多样性的理解，特别关注的是孑遗属 Chilelimnophila。柠檬科在化石记录中有详细记载，许多物种被认为是现代动物群中的孑遗物种。我们的研究基于在白垩纪西班牙和克钦琥珀中发现的第一批智利林虫化石标本。新材料允许建立一个新的Chilelimnophila (Ribeironia) Krzemiński、Kania-Kłosok et Arillo亚属。十一月以及三个新种：Chilelimnophila (Ribeironia) amorimi Krzemiński、Kania-Kłosok et Arillo, sp。十一月，Chilelimnophila（Chilelimnophila）wangi Krzemiński，Kania-Kłosok et Arillo，sp。 11 月，和 Chilelimnophila (Chilelimnophila) parva Krzemiński, Kania-Kłosok et Arillo, sp。十一月这项研究还显着扩展了该属的地层范围，提供了其存在可以追溯到白垩纪早期的证据。对 Limnophila 亚科 Limnophilinae 内的 Chilelimnophila 和相关属的形态和系统发育分析为这些生物的进化关系提供了宝贵的见解。此外，我们的研究强调了白垩纪西班牙和克钦琥珀地层的独特环境条件，这与这些生物体目前的栖息地有很大不同。