Representatives of the subfamily Cylapinae (Insecta: Heteroptera: Miridae) mostly inhabit the subtropics and tropics. The group currently comprises five tribes: Bothriomirini, Cylapini, Fulviini, Rhinomirini, and Vanniini. Additionally, the Psallopinae are sometimes included in this subfamily as a tribe. The interrelationships within Cylapinae are unknown, the monophyly is doubtful, and a phylogenetic study focusing on the entire subfamily has never been published. The molecular and fossil data analyses show that Miridae occurred at least in the Jurassic. The divergence dates for Cyalpinae tribes are unknown. We performed the phylogenetic analysis of Cylapinae based on molecular and morphological data, in addition to a combined dataset. We calibrated the molecular trees with fossil data. The results suggested non-monophyly of Cylapinae. The monophyly of Bothriomirini was confirmed. Cylapini were rendered as non-monophyletic. The Cylapus complex formed a clade with Vanniini, and the monophyly of Vanniini was doubtful. Fulviini were recovered as non-monophyletic, and the representatives fell into two separate well-supported clades. Dating analyses showed that the groups of the suprageneric level distributed across different continents diverged near the boarder between Mesosoic and Cenosoic, that was also true for the Asian suprageneric groups and the South American Cylapus complex. Australian groups of genera diverged in the Cenozoic.

中文翻译：

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Cylapinae 亚科的全证据系统发育及其亚群的分歧日期（昆虫纲：异翅目：Miridae）

Cylapinae亚科（昆虫纲：异翅目：Miridae）的代表大多栖息于亚热带和热带地区。该群体目前由五个部落组成：Bothriomirini、Cylapini、Fulviini、Rhinomirini 和 Vanniini。此外，Psallopinae 有时作为一个部落包含在该亚科中。Cylapinae 内部的相互关系尚不清楚，单系性值得怀疑，并且针对整个亚科的系统发育研究从未发表过。分子和化石数据分析表明，Miridae至少出现在侏罗纪。Cyalpinae 部落的分化日期尚不清楚。除了组合数据集之外，我们还根据分子和形态学数据对 Cylapinae 进行了系统发育分析。我们用化石数据校准了分子树。结果表明 Cylapinae 是非单系的。证实了 Bothriomirini 的单系性。Cylapini 被认为是非单系的。Cylapus复合体与Vanniini形成一个分支，Vanniini的单系性值得怀疑。Fulviini 被发现为非单系，其代表分为两个独立的、得到充分支持的进化枝。年代分析表明，分布于不同大陆的属上级类群在中生代和新生代边界附近出现分歧，亚洲的属上类群和南美洲的Cylapus杂岩也是如此。澳大利亚的属群在新生代出现分化。