Ichthyological Research is an official journal of the Ichthyological Society of Japan and is published quarterly in January, April, July, and November. Ichthyological Research primarily publishes research papers on original work, either descriptive or experimental, that advances the understanding of the diversity of fishes. Ichthyological Research strives to cover all aspects of fish biology, including taxonomy, systematics, evolution, biogeography, ecology, ethology, genetics, morphology, and physiology.

Ichthyological Research also publishes timely reviews and commentary on recent progress in active areas of research on fish biology.

Yoshiaki Kai 
Maizuru Fisheries Research Station
Field Science Education and Research Center, Kyoto University
Nagahama, Maizuru
Kyoto 625-0086, Japan
e-mail: mebaru@kais.kyoto-u.ac.jp

Managing Editor
Gen Kume
Faculty of Fisheries
Kagoshima University
4-50-20 Shimoarata, Kagoshima 890-0056, Japan
e-mail: kume@fish.kagoshima-u.ac.jp

Naoki Yagishita
Graduate School of Fisheries and
Environmental Sciences, Nagasaki
University, 1-14 Bunkyo-machi,
Nagasaki 852-8521, Japan
e-mail: n-yagi@nagasaki-u.ac.jp

Editors
Chris Burridge, University of Tasmania, Australia
Terry J. Donaldson, University of Guam, USA 
Yasuyuki Hashiguchi, Osaka Medical College, Japan
Yusuke Hibino, Kitakyushu Museum of Natural History and Human History, Japan 
Nao Kagawa, Kindai University, Japan
Mari Kawaguchi, Sophia University, Japan
Toshio Kawai, Hokkaido University, Japan 
Itsuro Koizumi, Hokkaido University, Japan
Tomoyuki Kokita, Fukui Prefectural University, Japan
B. Mabel Manjaji-Matsumoto, Universiti Malaysia Sabah, Malaysia
Mizuki Matsunuma, Kindai University, Japan
Yohei Nakamura, Kochi University, Japan
Makoto Okamoto, Marine Fisheries Research and Development Center (JAMARC), Japan
Yoshitaka Sakakura, Nagasaki University, Japan 
Aya Sato, Gunma University, Japan
Tamaki Shimose, Seikai National Fisheries Research Institute, Japan
Daisuke Tahara, Fukui Prefectural University, Japan
Hiroshi Takahashi, National Fisheries University, Japan
Tomohiro Takeyama, Okayama University of Science, Japan 
Yoshinori Taniguchi, Meijo University, Japan
Fumihito Tashiro, Hokkaido University, Japan
Thomas Trnski, Auckland Museum, New Zealand
William T. White, CSIRO, Australia

Advisory Board 
Jason B. Dunham, USA
Lee-Shing Fang, Taiwan
Graham S. Hardy, New Zealand
Jeffrey M. Leis, Australia
Keiichi Matsuura, Japan
Philip Munday, Australia
Edward O. Murdy, USA
Mutsumi Nishida, Japan
Lynne R. Parenti, USA
Mitsuhiko Sano, Japan
Kwang-Tsao Shao, Taiwan
Valentina G. Sideleva, Russia
Edward O. Wiley, USA