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Education: 1973 B. S. (Mathematics), National Tsing Hua University, Taiwan 1977 Ph.D. (Statistics), University of Wisconsin-Madison, USA Experiences: 1977-78 Visiting Assistant Professor, Department of Statistics University of Michigan-Ann Arbor, USA 1978-82 Associate Professor, National Tsing Hua University 1985-86 Visiting Scientist, Department of Biostatistics, Harvard University, USA 1982-present Professor, National Tsing Hua University Major Professional Services: 1994-2013 Associate Editor, Biometrics 1998-2011 Associate Editor, Journal of Agricultural, Biological, and Environmental Statistics 1991-96, 1999-2005 Associate Editor, Statistica Sinica 2001-2009 Associate Editor, Australian & New Zealand Journal of Statistics 2015- Associate Editor, Methods in Ecology and Evolution Referees for various journals in biology, ecology, statistics, epidemiology and related sciences. Honors: 1990-95 Taiwan National Science Council Award for Outstanding Research (傑出研究獎三次) 1987 Elected Member of International Statistical Institute 1996 Best Teaching Award, National Tsing Hua University (傑出教學獎) 1997 Fellow, Institute of Mathematical Statistics 2001 ISI Citation Classic Award for two highly-cited papers (Chao, 1984; Chao and Lee, 1992) 2003 Academic Award, Ministry of Education, Taiwan (教育部學術獎) 2005-2008 Taiwan National Chair Professor (國家講座) 2006- Tsing Hua Distinguished Chair Professor (清華特聘講座) Theses Supervised: ~ 100 Master Theses; 22 Ph.D. Theses Ph.D Theses: Lee, S-M (1990) Estimating population size for closed capture-recapture models via sample coverage. (Feng-Chia University) Ma, M-C (1992) Stopping rule and estimation for recapture debugging with unequal detection rates. (National Cheng-Kung University) Hwang, W. D. (1992) Quantifying the effects of unequal catchabilities on Jolly-Seber estimators. (National Cheng-Kung University) Chu, W-T (1995) A unified martingale estimation approach to estimating parameters of time-specific model. (Feng-Chia University) Chen, Y-C (1996) The estimation of population size for capture-recapture experiments under Pollock’s robust design. (Chia-Nan College of Pharmacy and Science) Hwang, Wen-Han (1997) Estimating population Size for continuous time capture-recapture experiments. (National Chung-Hsin University) Tsay, P-K (1997) Comparisons of loglinear model and sample coverage methods in capture-recapture experiments. (Chang-Kang University) Lin, S-P (2001) Comparison of population size estimation between continuous and discrete capture-recapture experiments. (Chung-Hua University) Yang, H-C (2002) The applications of Markov chain models and kernel smoothing in capture-recapture experiments. (Institute of Statistical Science, Academia Sinica) Shen, T.- J. (2003) Prediction of Biodiversity. (National Chung-Hsin University) Li, P.-C. (2005) Species estimation and similarity indices in quadrat sampling. (Wistron NeWeb Co. Industrial Park, Hsin-Chu) (Co-advisee) Lee, P.-L. (2006) Clustering for longitudinal data. (Department of Statistics, Tamkang University) Chiang, S.-C. (2006) Similarity indices for multiple communities. (Biostatistics Task Force, Taipei Veterans General Hospital) Pan, H. Y. (2007) Estimation of shared species richess in multiple communities. (National Chia-Yi University) Han, R.-P. (2008) Statistical estimation of beta diversity. (Central Taiwan University of Science and Technology) Jiang, Y.-H. (2008) Multiple-community similarity indices under both discrete and continuous diversity data. Lin, C.-W. (2009) Estimation of biodiversity in sampling without replacement. (Chaoyang University of Technology) Lin, S.-Y. (2012) Hierarchical and space-time decomposition of biodiversity indices. Wang, Y.-T. (2013) Statistical estimation of Hill numbers and similarity indices. Hsu, F.-J. (2013) Statistical estimation of rarefaction and extrapolation curve for beta diversity. Hsieh, T.-C. (2014) Rarefaction and extrapolation with biodiversity. Ma, K.-H. (2016) Rarefaction and extrapolation of similarity/differentiation measures and species diversity in sampling without replacement. Major Research Area: Ecological statistics and bio-diversity estimation •Species estimation and population size estimation •Animal abundance models and estimation •Capture-recapture models and their applications •Bio-diversity indices (species, phylogenetic and functional diversities) and their estimation •Biological sampling methods Citations and Recent Plenary talks: A full list of publications is given on the website. The total number of citations by SCI authors in Web of Science > 12600 citations; The total number of citations in Google Scholar > 23000 citations (Accessed on November, 2019) (1) Plenary speaker, 2016 British Ecological Society Annual Meeting, December, 2016, Liverpool, UK. (2) Plenary speaker, 2018 Australian Ecological Society Annual Conference, November 2018, Brisbane, Australia. (3) Plenary speaker, 2019 International Congress of Conservation Biology (ICCB), July, 2019, Kuala lumpur, Malaysia.

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1 Chao, A.(1981). Approximate mean squared errors of estimators reliability for k-out-of-m systems in the independent exponential case. Journal of American Statistical Association, 76, 720-724. (pdf) 2 Chao, A.(1981). On estimating the probability of discovering a new species. Annals of Statistics, 9, 1339-1342. (pdf) 3 Chao, A.(1982). On comparing estimators of P{Y.LT.X} in the exponential case. IEEE Transacations on Reliability, R-31, 389-392. (pdf) 4 Chao, A.(1982). Another approach to incomplete integrals. The American Statistician, 36, p.48. 5 Chao, A.(1983). Approximate mean squared errors of estimators of reliability in the two-parameter exponential case. Communications n Statistics, A12, 633-644. 6 Chao, A. and Hwang, W. D.(1983). Bayes estimation of reliability for special k-out-of-m:G systems. IEEE Transactions on Reliability, R-33, 246-249. (pdf) 7 Chao, A. (1984). Nonparametric estimation of the number of classes in a population. Scandinavian Journal of Statistics, 11, 265-270. (pdf) 8 Chao, A., Lin, S. and Lin, J. J. (1984). Jackknife estimator of an exponential parameter under type I censoring. Proceedings of National Science Council, 8, 191-194. 9 Cheng, K. F. and Chao, A. (1984). Confidence intervals for reliability from stress-strength relationships. IEEE Transactions on Reliability, R-33, 246-249. (pdf) 10 Chao, A. (1985). Asymptotic moments and normality of an estimator arising in life testing. Statistical Theory and Data Analysis (Matusita Ed.), North Holland, 153-158. 11 Chao, A. and Cheng, K. F. (1985). Interval estimation of reliability for highly reliable stress-strength models (In memory of Professor Cheng-Tan Hsiao). Chinese Journal of Mathematics, 13, 131-136. 12 Chao, A. and Hwang, S-J. (1986). Comparison of confidence intervals for the parameters of the Weibull and extreme value distribution. IEEE Transactions on Reliability, R-35, 111-113. (pdf) 13 Chao, A. and Hwang, L-C. (1987). A modified Monte Carlo technique for confidence limits on system reliability using pass-fail data. IEEE Transactions on Reliability, R-36, 109-112. (pdf) 14 Chao, A. (1987). Estimating the population size for capture-recapture data with unequal catchability. Biometrics, 43, 783-791. (pdf) 15 Chao, A. (1988). Estimating animal abundance with capture frequency data. Journal of Wildlife Management 52(2), 295-300. (pdf) 16 Chao, A. (1988). Estimating population size in a generalized capture-recapture model with applications epidemiological data. Statistical Theory and Data Analysis II (Matusita Ed.) North Holland, 29-36. 17 Chao, A., Lee, S-M and Chen, T-C. (1988). A generalized Good’s nonparametric coverage estimator. Chinese Journal of Mathematics, 16, 189-199. (pdf) 18 Chao, A. (1989). Estimating population size for sparse data in capture-recapture experiments. Biometrics, 45, 427-438. (pdf) 19 Chao, A. and Wang, N.S.(1990). Logistic regression analysis for hospital data. Journal of Chinese Statistical Association, 28, 92-103. (in Chinese) (pdf) 20 Chao, A. and Lee, S-M.(1990). Estimating the number of unseen species with frequency counts. Chinese Journal of Mathematics, 18, 335-351. 21 Chao, A., Chang, S-H. and Tsung, Y.H. (1991). Sexing Dunlins of Kung-Du area by a statistical method. Journal of Chinese Association, 29, 131-143. (in Chinese) 22 Chao, A., Lee, S-M and Jeng, C-L. (1992). Estimating population size for capture-recapture data when capture probabilities vary by time and individual animal. Biometrics, 48, 201-216. (pdf)(pdf+appendix) 23 Chao, A. and Lee, S-M. (1992). Estimating the number of classes via sample coverage. Journal of American Statistical Association, 87, 210-217. (pdf) 24 Yang, M.C.K. and Chao, A. (1992). On optimal stopping rules in software reliability. Computing Science and Statistics, Proceedings of the 23rd Symposium on the Interface, E.M.Karamidas and S.M. Kaufman (Editors), 102-105. 25 Chao, A.and Lee, S-M. (1993). Estimating population size for continuous-time capture-recapture models via sample coverage. Biometrical Journal, 35, 29-45. (pdf) 26 Ma,M-C. and Chao,A.(1993). Generalized sample coverage with an application to Chinese poem. Statistica Sinica, 3, 19-34. (pdf) 27 Chao, A. and Ma, M-C. and Yang, M. C. K.(1993), Stopping rule and estimation for recapture debugging with unequal detection rates. Biometrika, 80, 193-201. (pdf) 28 Lin, H-S., Chao, A. and Lee, S-M. (1993). Consistency of an estimator of the number of species–in memory of Professor Wen-Chen Chen. Journal of Chinese Statistical Association, 31, 253-270. (in Chinese) (pdf) 29 Lee, S-M. and Chao, A.(1994). Estimating population size for closed capture-recapture models via sample coverage. Biometrics, 50, 88-97. (pdf) 30 Hsieh, T-T, Chang, T-C, Chiu, T-H, Hsu, J-J, and Chao, A.(1994), Growth discordancy, birth weight and neonatal adverse events in third trimester twin gestations. Gynecologic and Obstetric Observation, 38, 36-40. 31 Chao, A. (1994) Reply to population size estimation for sparse data. Biometrics, 50,303-305. 32 Hwang, W-D. and Chao, A.(1995). Quantifying the effects of unequal catchabilities on Jolly-Seber estimators via sample coverage. Biometrics 51, 128-141. (pdf) 33 Yang, M. C. K. and Chao, A. (1995). Reliability estimation and stopping rules for software testing based on repeated appearances. IEEE Transactions on Reliability 44, 315-321. (pdf) 34 Chen, Y-C, Hwang, W-H, Chao, A. and Kuo, C-Y (1995). Estimating the number of common species: Analysis of the number of common bird species in two river estuaries. Journal of Chinese Statistical Association, 33, 373-393. (pdf) 35 Chao, A., Yip, P. and Lin, H-S. (1996). Estimating the number of species via martingale estimating function. Statistica Sinica, 403-418. (pdf) 36 Yip, P. and Chao, A. (1996). Estimating population size from capture-recapture via sample coverage and estimating function in continuous time. Stochastic Models 12,17-35. 37 Ho, T-P, Chao, A. and Yip, P (1997). Seasonal variation in suicides re-examined: no sex difference in Hong Kong and Taiwan. Acta Psychiatrica Scandinavica 95, 26-31. (pdf) 38 Yip, P., Chao, A. and Ho, T-P (1998). A re-examination of seasonal variation in suicides in Australia and New Zealand. Journal of Affective Disorders 47, 141-150. (pdf) 39 Chao, A. and Tsay, P. K. (1998). A sample coverage approach to multiple-system estimation with application to census undercount. Journal of American Statistical Association 93, 283-293. (pdf) 40 Chao, A. (1998). Capture-recapture models. p. 482-486 in Encyclopedia of Biostatistics (P. Armitage and T. Colton, Editors), Wiley. (pdf) 41 Chao, A. and Chang, S-H. (1999). An estimating function approach to catch-effort models. Environmental and Ecological Statistics 6, 313-334. (pdf) 42 Chao, A., Chu, Wenten and Hsu, C-H. (2000). Capture-recapture when time and behavioral response affect capture probabilities. Biometrics, 56, 427-433. (pdf) 43 Chao, A., Hwang, W-H, Chen, Y-C., and Kuo C-Y. (2000). Estimating the number of shared species in two communities. Statistica Sinica 10, 227-246. (pdf) 44 Yip, P. S. F., Xi, L., Chao, A. and Hwang, W-H (2000). Estimating the population size with a behavioral response in capture-recapture experiment. Environmental and Ecological Statistics, 7, 405-414. (pdf) 45 Chao, A., Lin, S.-P., Yang, H.-C. and Yip, P. S. F. (2000).The analysis of Hong Kong Big Bird Race data for the year of 2000. Journal of Chinese Statistical Association, 38, 231-241. (In Chinese with English abstract) (pdf) 46 Yip, P. S. F., Chao, A. and Chiu, C. (2000). Seasonal variation in suicides: diminished or vanished? British Journal of Psychiatry 177, 366-369. (pdf) 47 Chao, A., Yip, P., Lee, S-M. and Chu, W-T. (2001). Population size estimation based on estimating functions for closed capture-recapture models. Journal of Statistical Planning and Inference 92, 213-232. (pdf) 48 Chao, A. (2001). An overview of closed capture-recapture models. Journal of Agricultural, Biological and Environmental Statistics 6, 158-175. (pdf) 49 Tsay, P. K. and Chao, A. (2001). Population size estimation for capture-recapture models with applications to epidemiological data. Journal of Applied Statistics 28, 25-36. (pdf) 50 Chao, A., Tsay, P. K., Lin, S.-H., Shau, W.- Y. and Chao, D.- Y. (2001). Tutorial in biostatistics: the applications of capture-recapture models to epidemiological data. Statistics in Medicine, 20, 3123-3157. This paper is also selected in Tutorials in Biostatistics: Volume 1, R. B. D’Agostino (Ed), Wiley, Chichester, 2004. (pdf) 51 Hwang, W-H, Chao, A. and Yip, P. S. F. (2002). Continuous-time capture-recapture models with time variation and behavioral response. Australian and New Zealand Journal of Statistics 44, 41-54. (pdf) 52 Huggins, R. and Chao, A.(2002). Asymptotic properties of an optimal estimating function approach to the analysis of mark recapture data. Communications in Statistics – Theory & Methods 31, 575-595. (pdf) 53 Hwang, W. H. and Chao, A.(2002). Continuous-time capture-recapture models with covariates. Statistica Sinica 12, 1115-1131. (pdf) 54 Chao, A. and Bunge, J. (2002). Estimating the number of species in a stochastic abundance model. Biometrics 58, 531-539. (pdf) 55 Huggins, R., Yang, H.-C., Chao, A. and Yip, P. S. F. (2003). Population size estimation using local sample coverage for open populations. Journal of Statistical Planning and Inference 69, 113, 699-714. (pdf) 56 Shen, T, -J., Chao, A and Lin, J.-F. (2003). Predicting the number of new species in a further taxonomic sampling. Ecology 84, 798-804. (pdf) 57 Yip, P. S. F., Wang, Y. and Chao, A.(2003). The application of capture-recapture methods in software reliability. H. Pham (Editor), Handbook of Reliability Engineering, Chapter 27, 493-510. Springer. 58 Chao, A., Yang H.-C. and Yip, P. S. F. (2003). The use of capture-recapture methodology in epidemiological surveillance. Y. Lu and J.-Q. Fang (eds) Advanced Medical Statistics. Chapter 19, 711-739. International Science Press, Singapore. 59 Chao, A. and Shen, T.-J. (2003). Nonparametric Estimation of Shannon’s index of diversity when there are unseen species. Environmental and Ecological Statistics 10, 429-443. (pdf) 60 Chao, A. Book review for Borchers, D. L., Buckland, S. T. and Zucchini, W. Estimating Animal Abundance: Closed Populations.Springer, London, 2002. Biometrics 59, 460-461. (pdf) 61 Ida, H., Boylan, S. A., Weigel, A. L., Smit-McBride Z., Chao, A., Gao, J. Buchoff, P., Wistow, G. and Hjelmeland, L. M. (2004) EST analysis of mouse retina and REP/choroid cDNA libraries. Molecular Vision 10, 439-444. (pdf) 62 Chao, A. and Shen, T.-J. (2004). Non-parametric prediction in species sampling. Journal of Agricultural, Biological and Environmental Statistics 9, 253-269. (pdf) 63 Chao, A. (2005). Species estimation and applications. Encyclopedia of Statistical Sciences, 2nd Edition, Vol. 12, 7907-7916, (N. Balakrishnan, C. B. Read and B. Vidakovic, eds) Wiley, New York. (pdf) 64 Chao, A. and Huggins, R. M. (2005). Classical closed population models. The Handbook of Capture-Recapture Methods Chapter 2, 22-35 (Manly, B., McDonald, T. and Amstrup, S., Editors), Princeton University Press. 65 Chao, A. and Huggins, R. M. (2005). Modern closed population models. The Handbook of Capture-Recapture Methods Chapter 4, 58-87 (Manly, B., McDonald, T. and Amstrup, S., Editors), Princeton University Press. 66 Yip, P. S. F., Lam, K. F., Lau, E., Chau, P. H., Tsang, K. W. and Chao, A. (2005). A comparison study of real time fatality rates: SARS in Hong Kong, Singapore, Taiwan, Toronto and Beijing, China. Journal of Royal Statistical Society, Ser. A. 168, 233-243. (pdf) 67 Chao, A., Chazdon, R. L., Colwell, R. K. and Shen, T.-J.(2005). A new statistical approach for assessing similarity of species composition with incidence and abundance data. Ecology Letters 8, 148-159. (pdf file) Spanish translation in pp. 85-96 of Halffter, G. Soberon, J., Koleff, P. and Melic, A. (eds) 2005 Sobre Diversidad Biologica: el Sognificado de las Diversidades Alfa, Beta y Gamma. m3m-Monografias 3ercer Milenio, vol. 4, SEA, CONABIO, Grupo DIVERSITAS & CONACYT, Zaragoza. IV +242 pp. (pdf) 68 Yang H. C. and Chao, A. (2005). Modeling animals’ behavioral response by Markov chain models for capture-recapture experiments. Biometrics 61, 1010-1017. (pdf) 69 Lin, S.-P. and Chao, A. (2005). Discrete-time vs. continuous-time capture-recapture models-In honor of Professor T. Y. Hwang’s retirement. Journal of Chinese Statistical Association 43, 199-215. (pdf) 70 Chao, A., Chazdon, R. L., Colwell, R. K. and Shen, T.-J.(2006). Abundance-based similarity indices and their estimation when there are unseen species in samples. Biometrics 62, 361-371. (pdf) 71 Chao, A., Shen, T.-J. and Hwang, W.-H.(2006). Application of Laplace’s boundary-mode approximations to estimate species and shared species richness. Australian and New Zealand Journal of Statistics 48, 117-128. (pdf) 72 Chao, A., Li, P. C., Agatha, S. and Foissner W. (2006). A statistical approach to estimate soil ciliate diversity and distribution based on data from five continents. Oikos 114(3), 479-493. (pdf+appendix) 73 Liu, K., Beautrais, A., Caine, E., Chan, C., Chao, A., Conwell, Y., Law, C., Lee, D., Li, P. C., and Yip P. (2007) Charcoal burning suicides in Hong Kong and urban Taiwan ¡V an illustration of the impact of a novel method of suicide on overall regional rate.Journal of Epidemiology and Community Health 61, 248-253. (pdf) 74 Lo, T. C. T., Jiang, Y-H, Chao, A. and Chang, C. A. (2007) Use of statistical methods to find the polysaccharide structural characteristics and the relationships between monosaccharide composition ratio and macrophage stimulatory activity of regionally different strains of Lentinula edodes. Analytica Chimica Acta 584, 50-56. (pdf) 75 Yang, M. C. K., Chao, A. and Chen, Y. C. (2007). Assessing software reliability by unrevealed proportion estimation in stratified sampling. Journal of Chinese Statistical Association 45, 18-37. (pdf+appendix) 76 Foissner, W., Chao, A. and Katz L. A. (2008) Diversity and geographic distribution of ciliates (Protista: Ciliophora). Biodiversity and Conservation 17, 345-363. (pdf) 77 Chao, A., Jost, L., Chiang, S. C. Jiang Y.- H. and Chazdon, R. (2008). A Two-stage probabilistic approach to multiple-community similarity indices. Biometrics 64, 1178-1186. (pdf) 78 Chao, A., Law, C. K., Li, P. C. and Yip, P. S. F. (2008). Suicides in Taiwan, Chapter 6, 61-79 in Suicide in Asia: Causes and Prevention (Yip, P. S. F., Editor) University of Hong Kong Press. 79 Chao, A., Pan, H. Y. and Chiang, S. C. (2008). The Petersen-Lincoln estimator and its extension to estimate the size of a shared population. Biometrical Journal 50, 957-970. (pdf) 80 Chao, A., Colwell, R. K., Lin, C. W. and Gotelli, N. J. (2009). Sufficient sampling for asymptotic minimum species richness estimators. Ecology 90, 1125-1133. (pdf) (pdf+appendix) 81 Gossner, M., Chao, A., Bailey, R. and Prinzing, A. (2009). Native fauna on exotic trees: Phylogenetic conservatism and geographic contingency in two lineages of phytophages on two lineages of trees. The American Naturalist 173, 599-614. (pdf) (pdf+appendix) 82 Norden, N., Chazdon, R., Chao, A., Jiang, Y.-H. and Vilchez-Alvarado, B. (2009) Resilience of tropical rain forests: rapid tree community reassembly in secondary forests. Ecology Letters 12, 385-394. (pdf) 83 Gotelli, N. J., Anderson , M. J., Arita, H. T., Chao, A., Colwell, R. K., Connolly, S. R., Currie, D. J., Dunn, R. R., Graves, G. R., Green, J. L., Grytnes, J.-A., Jiang, Y.-H., Jetz , W., Lyons, S. K., McCain, C. M., Magurran, A. E., Rahbek, C., Rangel, T. F.L.V.B. , Soberon, J., Webb, C. O., Willig, M. R. (2009) Patterns and causes of species richness: a general simulation model for macroecology. Ecology Letters 12, 873-886. (pdf) 84 Pan, H. Y., Chao, A. and Foissner, W. (2009). A non-parametric lower bound for the number of species shared by multiple communities. Journal of Agricultural, Biological and Environmental Statistics. 14, 452-468. (pdf) 85 Jost, L., DeVries, P., Walla, T., Greeney, H., Chao, A. and Ricotta, C. (2010) Partitioning diversity for conservation analyses. Diversity and Distribution 16, 65-76. (pdf) 86 Chao, A., Chiu C.-H. and Jost, L. (2010). Phylogenetic diversity measures based on Hill numbers. Philosophical Transactions of the Royal Society B., 365, 3599-3609. (pdf) 87 Jost, L., Chao, A. and Chazdon, R. L. (2011). Compositional similarity and beta diversity. In Biological Diversity: Frontiers in Measurement and Assessment, pp. 66-84 (eds A. Magurran & B. McGill) Oxford: Oxford University Press. (pdf) 88 Chazdon, R. L., Chao, A., Colwell, R. K., Lin, S.-Y., Norden, N., Letcher, S. G., Clark, D. B., Finegan, B. and Arroyo J. P. (2011). A novel statistical method for classifying habitat generalists and specialists. Ecology, 92, 1332-1343. (pdf) (appendix) 89 Gotelli, N., Chao, A., Colwell, R. K., Hwang, W-H and Graves, G. R. (2012). Specimen-based modeling, stopping rules, and the extinction of the ivory-billed woodpecker. Conservation Biology 26, 47-56. (Cover Article) 89 (pdf) (pdf+appendix) 90 Colwell, R. K., Chao, A., Gotelli, N. J., Lin, S. Y., Mao, C. X., Chazdon, R. L.,and Longino, J. T. (2012). Models and estimators linking individual-based and sample-based rarefaction, extrapolation and comparison of assemblages. Journal of Plant Ecology 5, 3-21. (pdf) 91 Chao, A. and Jost, L. (2012). Diversity measures. In Encyclopedia of Theoretical Ecology (eds. A. Hastings and L. Gross), pp. 203-207, Berkeley: University of California Press. (pdf) 92 Chao, A. and Lin, C.-W. (2012). Nonparametric lower bound for species richness and shared species richness under sampling without replacement. Biometrics, 68, 912-921. (pdf)(pdf+appendix) 93 Chao, A., Chiu, C. H. and Hsieh, T. C. (2012) Proposing a resolution to debates on diversity partitioning. Ecology, 93, 2037-2051. (pdf) (pdf+appendix) 94 Eren, M. I., Chao, A., Hwang, W.-H. and Colwell, R. K. (2012). Estimating the richness of a population when the maximum number of classes is fixed: a nonparametric solution to an archaeological problem. PLoS One 7(5): e34179. (pdf) (pdf+appendix) 95 Chao, A. and Jost. L. (2012) Coverage-based rarefaction and extrapolation: standardizing samples by completeness rather than size. Ecology, 93, 2533-2547.(Cover Article) 95(pdf) (pdf+appendix) 96 Chao, A. and Chiu, C. H. (2012) Estimation of species richness and shared species richness. In N. Balakrishnan (Ed). Methods and Applications of Statistics in the Atmospheric and Earth Sciences. p.76-111, Wiley, New York. (pdf) 97 Gotelli, N. J., and Chao, A. (2013). Measuring and estimating species richness, species diversity, and biotic similarity from sampling data. In Levin, S. A. (Ed). Encyclopedia of Biodiversity, 2nd Edition, Vol. 5, pp. 195-211, Waltham, MA: Academic Press. (pdf) 98 Dornelas M., Magurran, A. E., Buckland, S. T., Chao, A., Chazdon, R. L., Colwell, R. K., Curtis, T., Gaston, K. J., Gotelli, N. J., Kosnik, M. A., McGill, B., McCune, J. L., Morlon, H., Mumby, P. J., Øvreås, L., Studeny, A., and Vellend, M. (2013). Quantifying temporal change in biodiversity: challenges and opportunities. Proceedings of the Royal Society B-Biological Sciences, 280, 20121931. (pdf) 99 Chao, A., Wang, Y. T., and Jost, L. (2013). Entropy and the species accumulation curve: a novel estimator of entropy via discovery rates of new species. Methods in Ecology and Evolution, 4, 1091-1110. (pdf) (pdf+appendix) (Rcode) 100 Müller, J., Bae, S., Röder, J., Chao, A. and Didham, R. K. (2014) Airborne LiDAR reveals context dependence in the effects of canopy architecture on arthropod diversity. Forest Ecology and Management, 312, 129-137. (pdf) (pdf+appendix) 101 Chiu, C.-H., Jost, L. and Chao*, A. (2014). Phylogenetic beta diversity, similarity, and differentiation measures based on Hill numbers. Ecological Monographs 84, 21-44. (pdf) (pdf+appendix) 102 Chao, A., Gotelli, N. G., Hsieh, T. C., Sander, E. L., Ma, K. H., Colwell, R. K. and Ellison, A. M. (2014). Rarefaction and extrapolation with Hill numbers: a framework for sampling and estimation in species biodiversity studies. Ecological Monographs 84, 45-67. (pdf) (pdf+appendix) (Main for iNEXT.r) (Subroutine for iNEXT.r) 103 Chiu, C.-H. and Chao, A. (2014). Distance-based functional diversity measures and their decomposition: a framework based on Hill numbers. PLoS ONE 9(7):e100014. (pdf) (pdf+appendix) (comment1) (comment2) (Rcode) 104 Chiu, C.-H., Wang Y. T., Walther B. A. and Chao, A. (2014). An improved non-parametric lower bound of species richness via a modified Good-Turing frequency formula. Biometrics, 70, 671-682. (pdf) (pdf+appendix) 105 Chao, A., Chiu, C.-H., and Jost, L. (2014). Unifying species diversity, phylogenetic diversity, functional diversity, and related similarity/differentiation measures through Hill numbers. Annual Reviews of Ecology, Evolution, and Systematics, 45, 297-324. (pdf) (pdf+appendix) 106 Chao, A., Chiu, C.-H., Hsieh, T. C., Davis, T., Nipperess, D., and Faith, D. (2015) Rarefaction and extrapolation of phylogenetic diversity. Methods in Ecology and Evolution, 6, 380-388. (Cover Article)(pdf) (pdf+appendix) 107 Chao, A., Hsieh, T. C., Chazdon, R. L., Colwell, R. K., and Gotelli, N. J. (2015). Unveiling the species-rank abundance distribution by generalizing the Good-Turing sample coverage theory. Ecology 96, 1189-1201. (pdf) (pdf+appendix) (R_code) 108 Chao, A., Jost, L., Hsieh, T.C., Ma, K. H., Sherwin, W. B. and Rollins, L. A. (2015). Expected Shannon entropy and Shannon differentiation between subpopulations for neutral genes under finite island model. PloS One 10(6):e0125471. (pdf) (pdf+appendix) (R_code)(Corrections) 109 Chao, A. and Jost, L. (2015). Estimating diversity and entropy profiles via discovery rates of new species. Methods in Ecology and Evolution, 6, 873-882. (pdf) (pdf+appendix) (R_code) 110 Chao, A. (2015). Capture-recapture for human populations. Wiley StatsRef: Statistics Reference Online. 1–16.DOI:10.1002/9781118445112.stat04855.pub2. (pdf) 111 Chao, A., Hsieh T. C. and Yang H. C. (2015). The use of capture-recapture methodology in epidemiological surveillance and ecological surveys. pp. 425-467 in Y. Lu and J.-Q. Fang, L. Tian and H. Jin (eds) Advanced Medical Statistics. 2nd Edition. World Scientific Publishing Co., Singapore. (pdf) 112 Eren, M. I., Chao, A., Chiu, C-H., Colwell, R. K., Buchanan, B., Boulanger, M. T., Darwent, J., and O’Brien, M. J. (2016) Statistical analysis of paradigmatic class richness supports greater paleoindian projectile-point diversity in the Southeast. American Antiquity, 81, 174-192. (pdf) (pdf+appendix) 113 Chiu, C-H. and Chao, A. (2016). Estimating and comparing microbial diversity in the presence of sequencing errors. PeerJ 4:e1634. (pdf) (pdf+appendix) (R_code) 114 Chao, A. and Chiu, C. H. (2016). Nonparametric estimation and comparison of species richness. Wiley Online Reference in the Life Sciences. In: eLS. John Wiley & Sons, Ltd: Chichester. DOI: 10.1002/9780470015902.a0026329. (pdf) 115 Chao, A., Chiu C.-H. and Jost L. (2016). Phylogenetic diversity measures and their decomposition: a framework based on Hill numbers. pp. 141-172 in Pellens R. and Grandcolas P. (eds) Biodiversity Conservation and Phylogenetic Systematics: Preserving our Evolutionary Heritage in an Extinction Crisis, Springer. (pdf) 116 Chao, A., Chiu C.-H. and Jost L. (2016). Commentary: Statistical challenges of evaluating diversity patterns across environmental gradients in mega-diverse communities. Journal of Vegetation Science 27, 437-438. (pdf) (pdf+appendix) 117 Chang, Y. H., Kuan, H. C., Hsieh, T. C., Ma, K. H., Yang, C. H., Hsu, W. B., Tsai, S. F., Chao, A. and Liu, H. H. (2016). Network signatures of IgG immune repertoires in hepatitis B associated chronic infection and vaccination responses. Scientific Reports, 6:26556. (pdf) (pdf+appendix) 118 Chao, A., and Chiu, C.-H. (2016). Bridging the variance and diversity decomposition approaches to beta diversity via similarity and differentiation measures. Methods in Ecology and Evolution, 7, 919-928. (Cover Article for Biogeography Virtual Issue)(pdf) (pdf+appendix) (Shiny App) 119 Chao, A. and Chiu, C. H. (2016). Species richness: estimation and comparison. Wiley StatsRef: Statistics Reference Online. 1-26. (pdf) 120 T.C. Hsieh, K. H. Ma, and Chao, A. (2016). iNEXT: An R package for rarefaction and extrapolation of species diversity (Hill numbers). Methods in Ecology and Evolution, 7, 1451-1456. (Cover Article) (pdf) (pdf+appendix). 121 Hsieh, T.C. and Chao, A. (2017). Rarefaction and extrapolation: making fair comparison of abundance-sensitive phylogenetic diversity among multiple assemblages. Systematic Biology 66, 100-111. (pdf) (pdf+appendix) 122 Chao, A., Colwell, R. K., Chiu, C.-H. and Townsend, D. (2017). Seen once or more than once: Using Good-Turing theory to estimate species richness using only unique observations and a species list. Methods in Ecology and Evolution. Cover Article for Statistical Ecology Virtual Issue.(pdf) (pdf+appendix) 123 Buchana, B., Chao, A., Chiu C, H., Colwell, R. K., O’Brien M. J., Werner, A. and Eren, M. I. (2017). Environment-induced changes in selective constraints on social learning during the peopling of the Americas. Scientific Reports, 7:44431. (pdf) 124 Chao, A. and Colwell, R.K. (2017). Thirty years of progeny from Chao’s inequality: estimating and comparing richness with incidence data and incomplete sampling (invited article). Statistics and Operation Research Transactions, 41, 3-54. (pdf) 125 Norden, N., Boukili, B., Chao, A., Ma, K. H., Letcher, S. G., and Chazdon, R. L. (2017). Opposing mechanisms affect taxonomic convergence between tree assemblages during tropical forest succession. Ecology Letters, 20, 1448-1458.(pdf)(pdf+appendix) 126 Chao, A., Chiu, C.-H., Colwell, R. K., Magnago, L. F. S., Chazdon, R. L. , and Gotelli, N. J. (2017). Deciphering the enigma of undetected species, phylogenetic, and functional diversity based on Good-Turing theory. Ecology, 98, 2914-2929. (pdf)(pdf+appendix) 127 Sherwin, W. B., Chao, A., Jost, L. and Smouse, P. E. (2017) Information theory broadens the spectrum of molecular ecology and evolution. Trends in Ecology and Evolution, 32, 948-963.(pdf)(pdf+appendix) 128 Gaggiotti, O. E., Chao, A., Peres-Neto, P., Chiu, C.-H., Edwards, C., Fortin, M.-J., Jost, L., Richards, C. M., and Selkoe, K. A. (2018). Diversity from genes to ecosystems: a unifying framework to study variation across biological metrics and scales. Evolutionary Applications, 11, 1176-1193. (pdf) (pdf+appendix) 129 Chao, A., Chiu, C.-H., Villéger, S., Sun, I.-F., Thorn, S., Lin, Y.-C., Chiang, J. M. and Sherwin, W. B. (2019). An attribute-diversity approach to functional diversity, functional beta diversity, and related (dis)similarity measures. Ecological Monographs, 89, e01343. 10.1002/ecm.1343. (pdf)(pdf+appendix) 130 Chao, A., Chiu, C.-H., Wu, S.-H., Huang, C. L. and Lin, Y.-C. (2019). Comparing two classes of species alpha diversities and their corresponding beta and (dis)similarity measures, with an application to the assessment of a reintroduction program of Formosan sika deer (Cervus nippon taiouanus). Methods in Ecology and Evolution,10, 1286-1297. (pdf)(pdf+appendix) 131 Huber, B. and Chao, A. (2019) Inferring global species richness from mega-transect data and undetected species estimates. Contributions to Zoology, 88, 42-53. (pdf)(pdf+appendix) 132 Chao, A., Colwell, R. K., Gotelli, N. J. and Thorn, S. (2019) Proportional mixture of two rarefaction/extrapolation curves to forecast biodiversity changes under landscape transformation. Ecology Letters 22, 1913-1922. (pdf)(pdf+appendix) 133 Chao, A. and Ricotta, C. (2019). Quantifying evenness and linking it to diversity, beta diversity, and similarity. Ecology, 100(12), e02852. (pdf)(pdf+appendix) 134 Rother, D. C., Liboni, A. P., Magnago, L. F. S., Chao, A., Chazdon, R. L. and Rodrigues, R. R. (2019). Ecological restoration increases conservation of taxonomic and functional beta diversity of woody plants in a tropical fragmented landscape. Forest Ecology and Management, 451, 117538. (pdf)(pdf+appendix) 135 Chao, A., Y. Kubota, D. Zelený, C.-H. Chiu, C.-F. Li, B. Kusumoto, M. Yasuhara, S. Thorn, C.-L. Wei, M. J. Costello, and R. K. Colwell (2020). Quantifying sample completeness and comparing diversities among assemblages. Ecological Research, 35, 292-314. (pdf)(pdf+appendix) 136 Thorn, S., Chao, A., Bernhardt-Römermann, M., Chen, Y-H, Georgiev, K. B., Heibl, C., Müller, J., Schäfer, H. and Bässler, C. (2020). Rare species, functional groups, and evolutionary lineages drive successional trajectories in disturbed forests. Ecology, 101, e02949. (pdf)(pdf+appendix) 137 Georgiev, K. B., Chao, A., Castro, J., Chen, Y-H, Choi, C-Y., Fontaine, J. B., Hutto, R. L., Lee, E-J., Müller, J., Rost, J., Żmihorski, M. and Thorn, S. (2020) Salvage logging changes the taxonomic, phylogenetic and functional successional trajectories of forest bird communities. Journal of Applied Ecology, 00:1-10. (pdf)(pdf+appendix) 138 Kusumoto, B., Costello, M. J., Kubota, Y., Shiono, T., Wei, C.-L., Yasuhara, M and Chao, A. (2020) Global distribution of coral diversity: biodiversity knowledge gradients related to spatial resolution. Ecological Research, 35, 315-326. (pdf)(pdf+appendix) 139 Colwell, R. K. and Chao, A. (2020) Measuring and comparing class diversity in archaeological assemblages: A brief guide to the history and state-of-the-art in diversity statistics. To appear as a chapter in Eren, M. I., and B. Buchanan, editors. Defining and measuring diversity in archaeology. 140 Bebber, M. R. and Chao, A. (2020) The diversity of North America’s “Old Copper” projectile points. To appear as a chapter in Eren, M. I., and B. Buchanan, editors. Defining and Measuring Diversity in Archaeology. 141 Müller, J., Ulyshen, M., Seibold, S. Cadotte, M., Chao, A., Bässler, C., Vogel, S., Stengel, E., Hagge, J., Weiß, I., Baldrian, P., Tláskal, V., and Thorn, S. (2020) Primary determinants of communities in deadwood vary among taxa but are regionally consistent. To appear in Oikos. 142 Thorn, S., Chao, A., Georgiev, K. B., Müller, J., Bässler, C., …, Leverkus, A. B. (2020) Estimating retention benchmarks for salvage logging to protect biodiversity. To appear in Nature Communications.

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